1 Richards DE, King KE, Tahar AA, Harberd NP. How gibberellin regulates plant growth and development: amolecular genetic analysis of gibberellin signaling [J]. Annu Rev Plant Physiol Plant Mol Biol, 2001, 52 (52): 67-88 2 黄先忠, 蒋才富, 廖立力, 傅向东. 赤霉素作用机理的分子基础与调控模式研究进展[J]. 植物学通报, 2006, 23 (5): 499-510 [Huang XZ, Jiang CF, Liao LL, Fu XD. Progress on molecular foundation of GA biosynthesis pathway and signaling [J]. Chin Bull Bot, 2006, 23 (5): 499-510] 3 Wen CK, Chang C. Arabidopsis RGL1 encodes a negative regulator of gibberellin responses [J]. Plant Cell Online, 2002, 14 (1): 87-100 4 Fu X, Sudhakar D, Peng J, Richards DE, Christou P, Harberd NP. Expression of Arabidopsis GAI in transgenic rice represses multiple gibberellin responses [J]. Plant Cell, 2011, 13: 1791-1802 5 王润青. 小麦DELLA获得性突变体矮变1号幼苗的抗盐和抗旱性研究[D]. 新乡: 河南科技学院, 2016 [Wang RQ. The studies on salt tolerance and drought resistance of a wheat DELLA gain-of-function mutant Aibian 1 seedling [D]. Xinxiang: Henan Institute of Science and Technology, 2016] 6 王利彬. 大豆苗期干旱和高温胁迫应答机制研究及关键转录因子的筛选[D]. 哈尔滨: 东北农业大学, 2018 [Wang LB. The study on response mechanism and screening of key factors under drought and high temperature stresses in Soybean (Glycine max (L.) Merrill) [D]. Harbin: Northeast Agricultural University, 2018] 7 Zhao B, Li H, Li J, Wang B, Dai C, Wang J, Liu K. Brassica napus DS-3, encoding a DELLA protein, negatively regulates stem elongation through gibberellin signaling pathway [J]. Theor Appl Genet, 2017, 130 (4): 727-741 8 吕西洋. 棉花DELLA蛋白GAI4功能鉴定[D]. 石河子: 石河子大学, 2013 [Lv XY. Functional identification of DELLA GAI4 gene in Gossypium hirsutum L. [D]. Shihezi: Shihezi University, 2013] 9 逯久幸. 梅花休眠相关DELLA基因的克隆与功能分析[D]. 北京: 北京林业大学, 2015 [Lu JX. Functional analysis of DELLA genes related to dormancy in Prunus mume [D]. Beijing: Beijing Forestry University, 2015] 10 卢艳芬, 卜芋芬, 郝素晓, 汪天宇, 姚允聪. 5种苹果属植物GAI基因的克隆及生物信息学分析[J]. 北京农学院学报, 2017, 32 (1): 1-6 [Lu YF, Bu YF, Hao SX, Wang TY, Yao YC. Cloning and bioinformatic analysis of GAI in the five Malus species [J]. J Beijing Univ Agric, 2017, 32 (1): 1-6] 11 刘杭, 杨贺忠, 李亮, 王芳, 吴少华, 李永裕. 梨休眠DELLA蛋白PpGAI基因的克隆与表达分析[J]. 分子植物育种, 2016, 14 (8): 1995-2002 [Liu H, Yang HZ, Li L, Wang F, Wu SH, Li YY. Cloning and expression analysis of PpGAI genes of DELLA protein related to dormancy from Pyrus [J]. Mol Plant Breed, 2016, 14 (8): 1995-2002] 12 Li WJ, Zhang JX, Sun HY, Wang SM, Chen KQ, Liu YX, Li H, Ma Y, Zhang ZH. FveRGA1, encoding a DELLA protein, negatively regulates runner production in Fragaria vesca [J]. Planta, 2018, 247 (4): 941-951 13 Peng JR, Richards DE, Hartley NM, Murphy GP, Devos KM, Flintham JE, Beales J, Fish LJ, Worland AJ, Pelica F, Sudhakar D, Christou P, Snape JW, Gale MD, Harberd NP. ‘Green revolution’ genes encode mutant gibberellin response modulators [J]. Nature, 1999, 400 (6741): 256-261 14 Yamamoto Y, Hirai T, Yamamoto E, Kawamura M, Sato T, Kitano H, Matsuoka M, Ueguchi-Tanaka M. A Rice gid1 suppressor mutant reveals that gibberellin is not always required for interaction between its receptor, GID1, and DELLA proteins [J]. Plant Cell Online, 2010, 22 (11): 3589-3602 15 Yasumura Y, Crumpton TM, Fuentes S, Harberd NP. Step-by-step acquisition of the gibberellin-DELLA growth-regulatory mechanism during land-plant evolution [J]. Curr Biol, 2007, 17 (14): 1225-1230 16 Ariizumi T, Lawrence PK, Steber CM. The role of two F-box proteins, SLEEPY1 and SNEEZY, in Arabidopsis GA signaling [J]. Plant Physiol, 2011, 155 (2): 765-775 17 Sun T, Gubler F. Molecular mechanism of gibberellins signalling in plants [J]. Annu Rev Plant Biol, 2004, 55 (1): 197-223 18 Itoh H, Ueguchi-Tanaka M, Sato Y, Ashikari M, Matsuoka M. The gibberellin signaling pathway is regulated by the appearance and disappearance of SLENDER RICE1 in Nuclei [J]. Plant Cell Online, 2002, 14 (1): 57-70 19 Fu X, Harberd NP. Auxin promotes Arabidopsis root growth by modulating gibberellin response [J]. Nature, 2003, 421 (6924): 740-743 20 张文颖, 王晨, 朱旭东, 马超, 王文然, 冷翔鹏, 郑婷, 房经贵. 葡萄全基因组DELLA蛋白基因家族鉴定及其应答外源赤霉素调控葡萄果实发育的特征[J]. 中国农业科学, 2018, 51 (16): 3130-3146 [Zhang WY, Zhu C, Zhu XD, Ma C, Wang WR, Leng XP, Zheng T , Fang JG. Genome-wide identification and expression of DELLA protein gene family during the development of grape berry induced by exogenous GA [J]. Sci Agric Sin, 2018, 51 (16): 3130-3146] 21 Lee KP, Piskurewicz U, Tureckova V, Strnadb M, Lopez-Molinaa L. A seed coat bedding assay shows that RGL2-dependent release of abscisic acid by the endosperm controls embryo growth in Arabidopsis dormant seeds [J]. PNAS, 2010, 107 (44): 19108-19113 22 Ueguchi TM, Matsuoka M. The perception of gibberellins: clues from receptor structure [J]. Curr Opin Plant Biol, 2010, 13 (5): 503-508 23 Cao D, Hussain A, Cheng H, Peng J. Loss of function of four DELLA genes leads to light- and gibberellin-independent seed germination in Arabidopsis [J]. Planta, 2005, 223 (1): 105-113 24 贾永鹏, 李开祥, 昝领兄, 姚艳梅, 杜德志. 甘蓝型油菜全基因组DELLA蛋白基因家族的鉴定和表达分析[J]. 中国油料作物学报, 2019, 41 (3): 360-368 [Jia YP, Li KX, Zan LX, Yao YM, Du DZ. Genome-wide identification and expression of DELLA protein family in Brassica napus [J]. Chin J Oil Crop Sci, 2019, 41 (3): 360-368] 25 陈宇杰, 刘飞, 梁菲菲, 石君, 赵焕梦. 蓖麻DELLA蛋白家族GAI基因克隆、表达及生物信息学分析[J]. 内蒙古民族大学学报 (自然科学版), 2017, 32 (4): 320-327 [Chen JY, Liu F, Liang FF, Shi J, Zhao HM. Cloning of castor DELLA protein family GAI gene expression and analysis of bioinformatics [J]. J Inner Mongolia Univ (Nat Sci Ed), 2017, 32 (4): 320-327] 26 Liao WB, Ruan MB, Cui BM, Xu NF, Lu JJ, Peng M. Isolation and characterization of a GAI/RGA-like gene from Gossypium hirsutum [J]. Plant Growth Regul, 2009, 58 (1): 35-45 27 温玮. 棉花DELLA蛋白的基因克隆与功能的初步鉴定[D]. 海口: 海南大学, 2010 [Wen W. Molecular cloning and functional characterizaiton of genes encoding a putative DELLA protein from cotton [D]. Haikou: Hainan University, 2010] 28 Li JY, Li CY, Smith SM. Hormone metabolism and signaling in plants [M]. Salt Lake: Academic Press, 2017: 107-160 29 Muangprom A, Thomas SG, Sun TP, Osborn TC. A novel dwarfing mutation in a green revolution gne from brassica rapa [J]. Plant Physiol, 2005, 137 (3): 931-938 30 Vandenbussche F, Fierro AC, Wiedemann G, Reski R, Straeten DV. Evolutionary conservation of plant gibberellin signalling pathway components [J]. BMC plant biol, 2007, 7 (1): 65 31 Gomi K, Sasaki A, Itoh H, Ueguchi-Tanaka M, Ashikari M, Kitano H, Matsuoka M. GID2, an F-box subunit of the SCF E3 complex, specifically interacts with phosphorylated SLR1 protein and regulates the gibberellin-dependent degradation of SLR1 in rice [J]. Plant J, 2004, 37 (4): 626-634 32 Dill A, Thomas SG, Hu J, Steber CM, Sun TP. The Arabidopsis F-Box protein SLEEPY1 targets gibberellin signaling repressors for gibberellin-induced degradation [J]. Plant Cell Online, 2004, 16 (6): 1392-1405 33 张新蕊. 生长素和赤霉素参与调节低磷胁迫下玉米根系形态的改变[D]. 济南: 山东大学, 2011 [Zhang XR. Auxin and gibberellins involved in the changes of maize root morphology during phosphorous starvation [D]. Jinan: Shandong University, 2011] 34 Fu XD, Richards DE, Ait-ali T, Hynes LW, Oughan H, Peng JR, Harberd NP. Gibberellin-mediated proteasome-dependent degradation of the barley DELLA protein SLN1 repressor [J]. Plant Cell Online, 2002, 14 (12): 3191-3200 35 Murase K, Hirano Y, Sun TP, Hakoshima T. Gibberellin-induced DELLA recognition by the gibberellin receptor GID1 [J]. Nature, 2008, 456 (7221): 459-463 36 Swain SM, Tseng T, Olszewski NE. Altered expression of SPINDLY affects gibberellin response and plant development [J]. Plant Physiol, 2001, 126 (3): 1174-1185 37 Ben-Nissan G, Cui W, Kim DJ, Yang YD, Yoo BC, Lee JY. Arabidopsis casein kinase 1-Like 6 contains a microtubule-binding domain and affects the organization of cortical microtubules [J]. Plant Physiol, 2008, 148 (4): 1897-1907 38 Zentella R, Hu J, Hsieh WP, Matsumoto PA, Dawdy A, Barnhill B, Oldenhof H, Hartweck LM, Maitra S, Thomas SG, Cockrell S, Boyce M, Shabanowitz J, Hunt DF, Olszewski NE, Sun TP. O-GlcNAcylation of master growth repressor DELLA by SECRET AGENT modulates multiple signaling pathways in Arabidopsis [J]. Gene Dev, 2016, 30 (2): 164-176 39 Qin Q, Wang W, Guo X, Yue J, Huang Y, Xu XF, Li J, Hou SW. Arabidopsis DELLA protein degradation is controlled by a Type-One protein phosphatase, TOPP4 [J]. Plos Genet, 2014, 10 (7): e1004464 40 Conti L, Nelis S, Zhang C, Woodcock A, Swarup R, Galbiati M, Tonelli C, Napier R, Hedden P, Bennett M, Sadanandom A. Small ubiquitin-like modifier protein SUMO enables plants to control growth independently of the phytohormone gibberellin [J]. Dev Cell, 2014, 28 (1): 102-110 41 Ariizumi T, Murase K, Sun TP, Steber SC. Proteolysis-independent downregulation of DELLA repression in Arabidopsis by the gibberellin receptor GIBBERELLIN INSENSITIVE DWARF1 [J]. Plant Cell, 2008, 20 (9): 2447-2459 42 郝鹏博. 外源赤霉素和多效唑对桃节间长度及赤霉素代谢基因表达影响[D]. 郑州: 河南农业大学, 2017 [Hao PB. Effects of GA3 and PBZ on internode length and gene expression of gibberellins metabolism in Peach (Prunus ersica L.) [D]. Zhengzhou: Henan Agricultural University, 2017] 43 任怡然, 赵强, 由春香, 郝玉金. 苹果MdRGL3a基因的功能分析[J]. 园艺学报, 2017, 44 (6): 1021-1028 [Ren YR, Zhao Q, You CX, Hao YJ. Function analysis of MdRGL3a gene in Malus x domestica [J]. Acta Horticul Sin, 2017, 44 (6): 1021-1028] 44 Gallego-Bartolomé J, Alabadí D, Blázquez MA. DELLA-induced early transcriptional changes during etiolated development in Arabidopsis thaliana [J]. PLoS ONE, 2011, 6 (8): e23918 45 Gallego-Bartolomé J, Arana MV, Vandenbussche F, Zádníková P, Minguet EG, Guardiola V, Straeten DV, Benkova E, Alabadí D, Blázquez MA. Hierarchy of hormone action controlling apical hook development in Arabidopsis [J]. Plant J, 2011, 67 (4): 622-634 46 Willige BC, Ogiso-Tanaka E, Zourelidou M, Schwechheimer C. WAG2 represses apical hook opening downstream from gibberellin and PHYTOCHROME INTERACTING FACTOR 5 [J]. Development, 2012, 139 (21): 4020-4028 47 Yanai O, Shani E, Dolezal K, Tarkowski P, Sablowski R, Sandberg G, Samach A, Ori N. Arabidopsis KNOXI proteins activate cytokinin biosynthesis [J]. Curr Biol, 2005, 15 (17): 1566-1571 48 Yaish MW, Ashraf EK, Tong Z, Beatty PH, Good AG, Bi YM, Rothstein SJ. The APETALA-2-Like transcription factor OsAP2-39 controls key interactions between abscisic acid and gibberellin in rice [J]. PLoS Genet, 2010, 6 (9): e1001098 49 Achard P, Cheng H, De GL. Integration of plant responses to environmentally activated phytohormonal signals [J]. Science, 2006, 311 (5757): 91-94 50 Zentella R, Zhang Z, Park M, Thomas SG, Endo A, Murase K, Fleet CM, Jikumaru Y, Nambara E, Kamiya Y. Global analysis of DELLA direct targets in early gibberellin signaling in Arabidopsis [J]. Plant Cell, 2007, 19 (10): 3037-3057 51 Ko J, Yang SH, Han K. Upregulation of an Arabidopsis RING-H2 gene, XERICO, confers drought tolerance through increased abscisic acid biosynthesis [J]. Plant J, 2006, 47 (3): 343-355 52 李志勇. 一个高秆显性四倍体小麦材料的发现与其DELLA蛋白丰度相关性分析[D]. 郑州: 河南农业大学, 2012 [Li ZY. Discovery of s high-stalk dominant mutant in tetraploid wheat and correlation analysis between its DELLA protein abundance and plant height [D]. Zhengzhou: Henan Agricultural University, 2012] 53 Achard P, Vriezen WH, Straeten D, Harberd NP. Ethylene regulates Arabidopsis development via the modulation of DELLA protein growth repressor function [J]. Plant Cell Online, 2003, 15 (12): 2816-2825 54 An F, Zhang X, Zhu Z. Coordinated regulation of apical hook development by gibberellins and ethylene in etiolated Arabidopsis seedlings [J]. Cell Res, 2012, 22 (5): 915-927 55 敖梦飞. 蜻蜓凤梨AfGAI基因的克隆与功能鉴定[D]. 海口: 海南大学, 2017 [Ao MF. Cloning and functional identification of AfGAI from Aechmea fasciata [D]. Haikou: Hainan University, 2017] 56 Luo J, Ma N, Pei H, Chen JW, Li J, Gao JP. A DELLA gene, RhGAI1, is a direct target of EIN3 and mediates ethylene-regulated rose petal cell expansion via repressing the expression of RhCesA2 [J]. J Exp Bot, 2013, 64 (16): 5075-5084 57 Zhou X, Zhang ZL, Park J, Tyler L, Yusuke J, Qiu K, Nam EA, Lumba S, Desveaux D, McCourt P, Kamiya Y, Sun TP. The ERF11 transcription factor promotes internode elongation by activating gibberellin biosynthesis and signaling [J]. Plant Physiol, 2016, 171 (4): 2760 58 Achard P, Baghour M, Chapple A, Hedden P, Van Der Straeten D, Genschik P, Moritz T, Harberd NP. The plant stress hormone ethylene controls floral transition via DELLA-dependent regulation of floral meristem-identity genes [J]. PNAS, 2007, 104 (15): 6484-6489 59 刘晔彤. 乙烯信号调控拟南芥发育时相转换的分子机制[D]. 上海: 上海师范大学, 2017 [Liu YT. Molecular mechanism of ethylene regulation in Arabidopsis thaliana phase transition [D]. Shanghai: Shanghai Normal University, 2017] 60 周峰. 植物防御反应及其光信号调控途径[J]. 北方园艺, 2015 (17): 179-182 [Zhou F. The plant defense response and its light signaling regulation pathways [J]. N Horticul, 2015 (17): 179-182] 61 Hou X, Lee L YC, Xia K, Yan Y, Yu H. DELLAs modulate jasmonate signaling via competitive binding to JAZs [J]. Dev Cell, 2010, 19 (6): 884-894 62 Bastian R, Dawe A, Meier S, Ludidi N, Bajic VB, Gehring C. Gibberellic acid and cGMP-dependent transcriptional regulation in Arabidopsis thaliana [J]. Plant Signal Behav, 2010, 5 (3): 224-232 63 Cheng H, Song S, Xiao L, Soo HM, Cheng Z, Xie D, Peng J. Gibberellin acts through jasmonate to control the expression of MYB21, MYB24, and MYB57 to promote stamen filament growth in Arabidopsis [J]. PLoS Genet, 2009, 5 (3): e1000440 64 吴绍华, 张世鑫, 陈月异, 田维敏. 橡胶树DELLA蛋白编码基因HbGAI的克隆与表达分析[J]. 西北植物学报, 2015, 35 (11): 2157-2163 [Wu SH, Zhang SX, Chen YY, Tian WM. Cloning and expression analysis of HbGAI gene in rubber tree (Hevea brasiliensis Muell. Arg.) [J]. Acta Bot Bor-Occident Sin, 2015, 35 (11): 2157-2163] 65 Navarro L, Bari R, Achard P. DELLAs control plant immune responses by modulating the balance of jasmonic acid and salicylic acid signaling [J]. Curr Biol, 2008, 18 (9): 650-655 66 Bai MY, Shang JX, Oh E, Fan M, Bai Y, Zentella R, Sun TP, Wang ZY. Brassinosteroid, gibberellin and phytochrome impinge on a common transcription module in Arabidopsis [J]. Nat Cell Biol, 2012, 14 (8): 810-817 67 王弋, 董晨, 魏永赞, 郑雪文, 李伟才. GA信号途径及其调控果树生长发育的研究进展[J]. 果树学报, 2018, 35 (4): 500-511 [Wang G, Wei RZ, Zheng XW, Li WC. Research progress on GA signaling pathway and its function in regulating fruit trees growth and development [J]. J Fruit Sci, 2018, 35 (4): 500-511] 68 Gallego-Bartolome J, Minguet EG, Grau-Enguix F, Abbas M, Locascio A, Thomas SG, Alabadí D, Blázquez MA. Molecular mechanism for the interaction between gibberllin and brassinosteroid signaling pathways in Arabidopsis [J]. PNAS, 2012, 109 (33): 13446-13451 69 Zhu J. Salt and drought stress signal transduction in plant [J]. Annu Rev Plant Biol, 2002, 53 (1): 247-273 70 边晓彤. DELLA家族在花生抗逆中的功能研究[D]. 济南: 山东师范大学, 2017 [Bian XT. Functionanl analysis of DELLA family proteins in Peanut stress resistance [D]. Jinan: Shandong Normal University, 2017] 71 Magome H, Yamaguchi S, Hanada A, Kamiya Y, Oda K. The DDF1 transcriptional activator upregulates expression of a gibberellin-deactivating gene, GA2ox7, under high-salinity stress in Arabidopsis [J]. Plant J, 2008, 56 (4): 613-626 72 张彤, 赵琳, 赵建刚, 李文滨. 植物DELLA蛋白的功能及其在大豆中的研究[J]. 大豆科学, 2011, 30 (5): 874-879 [Zhang T, Zhao L, Zhao G, Li WB. Function of plant DELLA protein and its research advancement in Soybean [J]. Soybean Sci, 2011, 30 (5): 874-879] 73 Wang KLC, Li H, Ecker JR. Ethylene biosynthesis and signaling networks [J]. Plant Cell, 2002, 14 (s1): S131-S151 74 王润青, 樊晓聪,宋梅芳, 肖阳, 郭林, 孟凡华, 杨青华, 吴大付, 杨建平. 小麦DELLA获得性突变体矮变1号增强了幼苗的抗盐能力[J]. 作物学报, 2016, 42 (11): 1721-1726 [Wang RQ, Fan XC, Song MF, Xiao Y, Guo L, Meng FD, Yang QH, Wu DF, Yang JP. A wheat DELLA gain-of-function mutant aibian 1 promotes seedling salt tolerance [J]. Acta Agron Sin, 2016, 42 (11): 1721-1726] 75 Pantin F, Monnet F, Jannaud D. The dual effect of abscisic acid on stomata [J]. New Phytol, 2012, 197 (1): 65-72 76 Acharya BR, Jeon BW, Zhang W, Assmann SM. Open Stomata 1 (OST1) is limiting in abscisic acid responses of Arabidopsis guard cells [J]. New Phytol, 2013, 200 (4): 1049-1063 77 Lin QB, Wu FQ, Sheng PK, Zhang Z, Zhang X, Guo XP, Wang JL, Cheng ZJ, Wang J, Wang HY, Wan JM. The SnRK2-APC/CTE regulatory module mediates the antagonistic action of gibberellic acid and abscisic acid pathways [J]. Nat Com, 2015, 6 (1): 7981 78 Nir I, Shohat H, Panizel I, Olszewski N, Aharoni A, Weiss D. The tomato DELLA protein PROCERA acts in guard cells to promote stomatal closure [J]. Plant Cell, 2018, 29 (12): 3186-3197 79 张涵, 王学敏, 刘希强, 马琳, 温红雨, 王赞. 紫花苜蓿MsGAI的克隆、表达及遗传转化[J]. 中国农业科学, 2019, 52 (2): 201-214 [Zhang H, Wang XM, Liu XQ, Ma L, Wen HY, Wang Z. Cloning expression analysis and transformation of MsGAI gene from Medicago sativa L. [J]. Sci Agric Sin, 2019, 52 (2): 201-214] 80 王玮, 管利萍, 张静, 侯岁稳. 拟南芥DELLA蛋白编码基因RGA和GAI的原核表达和多克隆抗体制备[J]. 兰州大学学报: 自然科学版, 2016, 52 (3): 422-428 [Wang W, Guan LP, Zhang J, Hou SW. Prokaryotic expression and polyclonal antibodies preparation of the DELLA protein-coding gene, RGA and GAI, in Arabidopsis [J]. J Lanzhou Univ (Nat Sci Ed), 2016, 52 (3): 422-428] 81 王玮, 冯起, 张莉环, 杨宁. DELLA蛋白缺失对拟南芥干旱胁迫耐受性的影响[J]. 西北植物学报, 2018, 38 (5): 867-872 [Wang W, Feng Q, Zhang LH, Yang N. Drought tolerance of DELLA proteins deficiency in Arabidopsis [J]. Acta Bot Boral-Occid Sin, 2018, 38 (5): 867-872] 82 朱珍. 赤霉素调控采后番茄果实抗冷机制研究[D]. 北京: 中国农业科学院, 2016 [Zhu Z. The mechanism of gibberellins in regulation of chilling tolerance of postharvest tomato fruit [D]. Beijing: Chinese Academy of Agricultural Sciences, 2016] 83 Djakovic-Petrovic T, de Wit M, Voesenek LA, Pierik R. DELLA protein function in growth responses to canopy signals [J]. Plant J, 2007, 51 (1): 117-126 84 Achard P, Liao L, Jiang C, Desnos T, Bartlett J, Fu X, Harberd NP. DELLAs contribute to plant photomorphogenesis [J]. Plant Physiol, 2007, 143 (3): 1163-1172 85 Feng S, Martinez C, Gusmaroli G, Wang Y, Zhou J, Wang F, Chen L, Yu L, Iglesias-Pedraz JM, Kircher S, Sch?fer E, Fu X, Fan LM, Deng XW. Coordinated regulation of Arabidopsis thaliana development by light and gibberellins [J]. Nature, 2008, 451 (7177): 475-479 86 Olsen JE. Light and temperature sensing and signaling in induction of bud dormancy in woody plants [J]. Plant Mol Biol, 2010, 73 (1-2): 37-47 87 欧阳芳群. 云杉幼苗生长光调控的生理与分子机制[D]. 北京: 北京林业大学, 2015 [Ouyang QF. Physiological and molecular mechanisms of light regulation on plant growth in spruce species [D]. Beijing: Beijing Forestry University, 2015] 88 Achard P, Renou J, Berthomé R, Harberd NP, Genschik P. Plant DELLAs restrain growth and promote survival of adversity by reducing the levels of reactive oxygen species [J]. Curr Biol, 2008, 18 (9): 656-660 89 Gapper C, Dolan L. Control of plant development by reactive oxygen species [J]. Plant Physiol, 2006, 141 (2): 341-345 90 Kwak JM, Mori IC, Pei ZM, Leonhardt N, Torres MA, Dangl JL, Bloom RE, Bodde S, Jones JD, Schroeder JI. NADPH oxidase AtrbohD and AtrbohF genes function in ROS-dependent ABA signaling in Arabidopsis [J]. EMBO J, 2003, 22 (11): 2623-2633 91 姚涛, 白素兰, 李苗苗, 张耀川, 何奕昆. DELLA蛋白参与拟南芥幼苗对一氧化氮逆境的抵抗[J]. 植物学报, 2011, 46 (5): 481-488 [Yao T, Su BL, Li MM, Zhang YC, He YK. DELLA protein is involved in the resistance of Arabidopsis seedlings to nitric oxide stress [J]. Chin Bull Bot, 2011, 46 (5): 481-488] 92 Jiang CF, Gao XH, Liao LL, Harberd NP, Fu XD. Phosphate starvation root architecture and anthocyanin accumulation responses are modulated by the gibberellin-DELLA signaling pathway in Arabidopsis [J]. Plant Physiol, 2007 , 145 (4): 1460-1470 93 Hauvermale AL, Ariizumi T, Steber CM. Gibberellin signaling: a theme and variations on DELLA repression [J]. Plant Physiol, 2012, 160 (1): 83-92 94 丁伟. 缺铁胁迫对梨氮代谢及赤霉素信号转导相关基因表达的影响[D]. 合肥: 安徽农业大学, 2015 [Ding W. Effects of iron-deficiency stress on the related genes expression of the nitrogen metabolism and GA signal transduction of pear [D]. Hefei: Anhui Agricultural University, 2015] 95 于璐. 芍药DELLA蛋白PlGAI1基因启动子的克隆及功能初步分析[D]. 沈阳: 沈阳农业大学, 2017 [Yu L. Cloning and functional identification of DELLA protein PlGAI1 genes promoter of the Paeonia lactiflora [D]. Shenyang: Shenyang Agricultural University, 2017] 96 Lim S, Park J, Lee N, Jeong J, Toh S, Watanabe A, Kim J, Kang H, Kim DH, Kawakami N, Choi G. ABA-insensitive3, ABA-insensitive5, and DELLAs interact to activate the expression of SOMNUS and other high-temperature-inducible genes in imbibed seeds in Arabidopsis [J]. Plant Cell, 2013, 25 (12): 4863-4878 97 白青松. 毛竹SAUR、DELLA基因的鉴定、克隆及功能分析[D]. 北京: 中国林业科学研究院, 2017 [Bai QS. Identification, clone and function analysis of SAUR and DELLA genes in Moso bamboo [D]. Beijing: Chinese Academy of Forestry, 2017] 98 Marín-de la Rosa N, Sotillo B, Miskolczi P, Gibbs DJ, Vicente J, Carbonero P, O?ate-Sánchez L, Holdsworth MJ, Bhalerao R, Alabadí D, Blázquez MA. Large-scale identification of gibberellin-related transcription factors defines group VII ethylene response factors as functional DELLA partners [J]. Plant Physiol, 2014, 166 (2): 1022-1032 99 Stewart Lilley JL, Gan Y, Graham IA, Nemhauser JL. The effects of DELLAs on growth change with developmental stage and brassinosteroid levels [J]. Plant J, 2013, 76 (1): 165-173 100 Oh E, Zhu JY, Bai MY. Cell elongation is regulated through a central circuit of interacting transcription factors in the llrabidoFasis hypocotyl [J]. Elife, 2014, 3 (327): 714-723 101 Weston DE, Elliott RC, Lester DR, Rameau C, Reid JB, Murfet IC, Ross JJ. The Pea DELLA proteins LA and CRY are important regulators of gibberellin synthesis and root growth [J]. Plant Physiol, 2008, 147 (1): 199-205 102 李秀峰. 黑龙江省水稻抽穗期基因的效应研究及育种应用[D]. 哈尔滨: 中国科学院大学, 2016 [Li XF. The study on effect of heading date genes in Heilongjiang Rice cultivars and their application in molecular assistant breeing [D]. Harbin: University of Chinese Academy of Sciences, 2016] 103 刘明明, 时振英, 张笑寒, 汪明璇, 张林, 郑克志, 刘继云, 胡兴明, 底翠茹, 钱前, 何祖华, 杨东雷. 调控IPA1培育高产高抗水稻新品种[C]. 上海: 第七届长三角植物科学研讨会暨青年学术报告会, 2018 [Liu MM, Shi ZY, Zhang XH, Wang MX, Zhang L, Zheng KZ, Liu JY, Hu XM, Di CR, Qian Q, He ZH, Yang DL. Regulating ipa1 to cultivate new rice varieties of high yield and high resistance [C]. Shanghai: The 7th Yangtze River Delta Plant Science Symposium and Youth Symposium, 2018] 104 麻楠, 刘静, 陈琦, 郝素晓,卜芋芬, 卢艳芬, 姚允聪. 苹果砧木‘SH6’中RGLs基因的克隆及生物信息学分析[J]. 北京农学院学报, 2018, 33 (2): 1-6 [Ma N, Liu J, Chen Q, Hao SX, Bu YF, Lu YF, Yao YC. Cloning and bioinformatic analysis of RGLs in apple rootstock‘SH6’[J]. J Beijing Univ Agric, 2018, 33 (2): 1-6] 105 刘超, 吴江生. 半矮秆基因BnaA6. rga-ds的亚细胞定位和转基因甘蓝型油菜的鉴定[J]. 中国油料作物学报, 2015, 37 (4): 427-432 [Liu C, Wu JS. Subcellular localization of semi-dwarf gene BnaA6. rga-ds and identification of its transgenic Brassica napus L. [J]. Chin Oil Crop Sci, 2015, 37 (4): 427-432] 106 潘现飞. 毛竹赤霉素信号途径相关基因的克隆与功能分析[D]. 杭州: 浙江农林大学, 2016 [Pan XF. Cloning and function analysis of genes related to gibberellin acid signaling in Phyllostachys edulis [D]. Hangzhou: Zhejiang A & F University, 2016] 107 黄昊. 珍珠黄杨DELLA蛋白家族BsGAI1, BsGAI2基因的克隆及功能分析[D]. 南京: 南京林业大学, 2017 [Huang H. Cloning and functional analysis of DELLA protein family BsGAI1 and BsGAI2 from Euonymus japonicus [D]. Nanjing: Nanjing Agricultural University, 2017] 108 田海霞. 拟南芥中茉莉素和赤霉素对WD-repeat/bHLH/MYB复合体的调节[D]. 长沙: 湖南农业大学, 2017 [Tian HX. Regulation of the WD-repeat/bHLH/MYB complex by gibberellin and jasmonate [D]. Changsha: Hunan Agricultural University, 2017] 109 Chen L, Xiang S, Chen Y, Li DB, Yu DQ. Arabidopsis WRKY45 interacts with the DELLA protein RGL1 to positively regulate age-triggered leaf senescence [J]. Mol Plant, 2017, 10 (9): 1174-1189 110 Gou JQ, Strauss SH, Tsai CJ, Fang K, Chen Y, Jiang X, Busov VB. Gibberellins regulate lateral root formation in Populus through interactions with auxin and other hormones [J]. Plant Cell, 2010, 22 (3): 623-639 111 李霞. DELLAs-MtDIP1-RAM1调控苜蓿-丛枝菌根共生的作用机制[D]. 开封: 河南大学, 2015 [Li X. The role of DELLAs-MtDIPl-RAM1 protein complex in arbuscular mycorrhizal symbiosis [D]. Kaifeng: Henan University, 2015] 112 樊玉倩, 肖爱芳, 张忠明. 百脉根起始结瘤相关转录因子间相互作用的鉴定[J]. 华中农业大学学报, 2019, 38 (1): 10-15 [Fan YQ, Xiao AF, Zhang ZM. Identification of interactions between transcription factors related to initial nodulation of Lotus japonicus [J]. J Huazhong Agric Univ, 2019, 38 (1): 10-15] 113 石玲玲, 王之, 李志英, 雷明, 徐立. 蜻蜓凤梨AfPIF4-1基因的克隆与遗传转化[J]. 分子植物育种, 2016, 14 (1): 66-71 [Shi LL, Wang Z, Li ZY, Lei M, Xu L. Cloning and genetic transformation of AfPIF4-1 in Aechmea fasciata [J]. Mol Plant Breed, 2016, 14 (1): 66-71] 114 宋晓波, 王红霞, 张志华. ‘辽宁2号’DELLA蛋白编码基因的克隆及表达分析[J]. 河北农业大学学报, 2016, 39 (1): 29-34 [Song XB, Wang HX, Zhang LH. Cloning and expression analysis of JrGAI gene in DELLA protein from‘Liaoning 2’ [J]. J Publish Dep Agric Univ Hebei, 2016, 39 (1): 29-34] 115 刘杭, 王芳, 李亮, 王文婷, 杨贺忠, 马鑫瑞, 吴少华, 李永裕. 沙梨PpGAI3基因的分离与原核表达分析[J]. 果树学报, 2017, 34 (2): 137-144 [Liu H, Wang F, Li L, Wang WT, Yang HZ, Ma XR, Wu SH, Li RY. Isolating and prokaryotic expression analysis of the PpGAI3 gene in Pyrus pyrifolia [J]. J Fruit Sci, 2017, 34 (2): 137-144] 116 吴红, 刘春英, 付祥梅, 盖树鹏, 张玉喜. 牡丹休眠相关基因PsGRASs筛选、克隆和表达特性分析[J]. 园艺学报, 2019, 46 (2): 365-374 [Wu H, Liu CY, Fu XM, Gai SP, Zhang YX. Screening, cloning and expression patterns analysis of PsGRASs associated with dormancy release in Tree Peony (Paeonia suffruticosa) [J]. Acta Horticul Sin, 2019, 46 (2): 365-374] 117 Martí C, Orzáez D, Ellul P, Moreno V, Carbonell J, Granell A. Silencing of DELLA induces facultative parthenocarpy in tomato fruits [J]. Plant J, 2007, 52 (5): 865-876 118 杨敏. 枇杷DELLA基因的克隆与分析[D]. 重庆: 西南大学, 2011 [Yang M. DELLA genes clone and analysis in Loquat (Eriobotrya japonica Lindl.) [D]. Chongqin: Southwest University, 2011] 119 张文颖, 朱旭东, 崔腾飞, 贾海锋, 房经贵, 王晨. 葡萄VvRGL基因应答GA调控无核果实发育的研究[J]. 西北植物学报, 2019, 39 (3): 381-390 [Zhang WY, Zhu XD, Cui TF, Jia HF, Fang JG, Wang C. VvRGL genes regulate seedless fruit development by responding to GA in Grapevine [J]. Acta Bot Boral-Occident Sin, 2019, 39 (3): 381-390] 120 Liu MY, Huang L, Ma ZT, Sun WJ, Wu Q, Tang ZZ, Bu TL, Li CL, Chen H. Genome-wide identification, expression analysis and functional study of the GRAS gene family in Tartary buckwheat (Fagopyrum tataricum) [J]. BMC Plant Biol, 2019, 19 (1): 342 121 夏小丛. 棉花(Gossypium hirsutum)GhJAZ3和GhGAI1在JA和GA介导的转基因拟南芥表皮毛分化和伸长中的功能研究[D]. 武汉: 华中师范大学, 2016 [Xia XC. Cotton (Gossypium hirsutum) GhJAZ3 and GhGAI1 functions in jasmonate and gibberellin mediated epidermal cell differentiation and elongation of transgenic Arabidopsis [D]. Wuhan: Central China Normal University, 2016] 122 De Vleesschauwer D, Seifi HS, Filipe O, Haeck A, Huu SN, Demeestere K, H?fte M. The DELLA protein SLR1 integrates and amplifies salicylic acid- and jasmonic acid-dependent innate immunity in Rice [J]. Plant Physiol, 2016, 170 (3): 1831-1847 123 Hosokawa K. Cell layer-specific accumulation of anthocyanins in response to gibberellic acid in tepals of Hyacinthus orientalis [J]. Biosci Biotechnol Biochem, 1999, 63 (5): 930-931 124 王可可. 外源赤霉素在心里美萝卜幼苗花青素代谢中作用研究[D]. 新乡: 河南师范大学, 2014 [Wang KK. The research of exogenous gibberellin treatment in anthocyanin metabolism in radish [D]. Xinxiang: Henan Normal University, 2014] 125 Xie Y, Tan HJ, Ma ZX, Huang J. DELLA proteins promote anthocyanin biosynthesis via sequestering MYBL2 and JAZ suppressors of the MYB/bHLH/WD40 complex in Arabidopsis thaliana [J]. Mol Plant, 2016, 9 (5): 711-721 126 Hatlestad GJ, Akhavan NA, Sunnadeniya RM, Elam L, Cargile S, Hembd A, Gonzalez A, McGrath JM, Lloyd AM. The beet Y locus encodes an anthocyanin MYB-like protein that activates the betalain red pigment pathway [J]. Nat Genet, 2014, 47 (1): 92-96 127 赵春丽, 王晓, 潘君飞, 彭丽云, 赖钟雄, 刘生财. 苋菜AtGAI基因克隆及表达分析[J]. 西北植物学报, 2019, 39 (2): 199-209 [Zhao CL, Wang X, Pan JF, Peng LY, Lai ZX, Liu SC. Cloning and expression analysis of AtGAI gene in Amaranthus tricolor L. [J]. Acta Bot Boral-Occid Sin, 2019, 39 (2): 199-209] 128 Cheminant S, Wild M, Bouvier F, Pelletier S, Renou JP, Erhardt M, Hayes S, Terry MJ, Genschik P, Achard P. DELLAs regulate chlorophyll and carotenoid biosynthesis to prevent photooxidative damage during seedling deetiolation in Arabidopsis [J]. Plant Cell, 2012, 23 (5): 1849-1860 129 周龙华. 甘蓝型油菜脂肪酸合成抑制与消减因素的研究[D]. 杭州: 浙江大学, 2016 [Zhou LH. Investigation on the inhibitors and fatty acid decomposers limiting seed oil accumulation in rapeseed (Brassica napus L.) [D]. Hangzhou: Zhejiang University, 2016]